Charlie Crist was a popular GOP governor of Florida. He will now run for the office as a Democrat.
Jason Thigpen, who would once qualify as a pretty middle of the road Republican, is going to run for Congress as a Democrat.
The question of what happens to moderates within the GOP is a fascinating one. There might be a place for moderate Republicans in certain northeastern areas of the country, because it will be hard for a Tea Party primary challenge to be successful. But in the South, the Plains and the Southwest, there simply isn't any tolerance by the GOP primary voters (ie Tea Party) for moderate Republicanism.
This is how a party becomes irrelevant.
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